[1] Kyungpook Natl Univ, Sch Earth Syst Sci, Dept Oceanog, Daegu, South Korea
[2] Kyungpook Natl Univ, Sch Earth Syst Sci, Kyungpook Inst Oceanog, Daegu, South Korea
[3] Dalhousie Univ, Dept Biol, Halifax, NS B3H 4R2, Canada
[4] Canadian Inst Adv Res, Program Integrated Microbial Divers, Toronto, ON, Canada