[1] Univ Toronto, Dept Ophthalmol & Vis Sci, Toronto, ON, Canada
[2] Univ Toronto, Mol Genet, Toronto, ON, Canada
[3] McMaster Univ, Div Ophthalmol, Hamilton, ON, Canada
[4] Univ Montreal, Ctr Rech, Maisonneuve Rosemont Hosp, Montreal, PQ, Canada
[5] Univ Toronto, Chem Engn & Appl Chem, Toronto, ON, Canada
[6] Toronto Western Hosp, Vis Sci Res Program, Toronto, ON, Canada