[1] South China Agr Univ, Coll Anim Sci, Lingnan Guangdong Lab Modern Agr, Guangzhou 510642, Peoples R China
[2] Minist Agr, Guangdong Prov Key Lab Agroanim Genom & Mol Breed, Guangzhou 510642, Peoples R China
[3] Minist Agr, Key Lab Chicken Genet Breeding & Reprod, Guangzhou 510642, Peoples R China
[4] Guangdong Engn Res Ctr Vector Vaccine Anim Virus, Guangzhou 510642, Peoples R China
[5] South China Collaborat Innovat Ctr Poultry Dis Co, Guangzhou 510642, Peoples R China
[6] ARS, Avian Dis & Oncol Lab, USDA, E Lansing, MI 48823 USA