[1] Univ Castilla La Mancha, Dept Ciencias Ambientales, E-45071 Toledo, Spain
[2] Univ Cantabria, CSIC, Inst Fis Cantabria, Grp Meteorol, E-39005 Santander, Spain
[3] Commiss European Communities, Joint Res Ctr, Inst Environm & Sustainabil, I-21027 Ispra Varese, Italy
[4] US Geol Survey, Sequoia Kings Canyon Field Stn, Western Ecol Res Ctr, Three Rivers, CA 93271 USA
[5] Univ Calif Los Angeles, Dept Ecol & Evolutionary Biol, Los Angeles, CA 90095 USA