[1] Uppsala Univ, Sci Life Lab, Dept Cell & Mol Biol, Uppsala, Sweden
[2] Oregon State Univ, Dept Microbiol, Corvallis, OR 97331 USA
[3] Uppsala Univ, Dept Med Biochem & Microbiol, Uppsala, Sweden
[4] Royal Netherlands Inst Sea Res, Dept Marine Microbiol & Biogeochem, NIOZ, NL-1790 AB Den Burg, Netherlands
[5] Univ Utrecht, NL-1790 AB Den Burg, Netherlands