[1] Univ Queensland, Sch Geog Planning & Environm Management, Brisbane, Qld 4072, Australia
[2] Arctic Univ Norway, Dept Arctic & Marine Biol, Tromso, Norway
[3] Jagiellonian Univ, Inst Environm Sci, PL-31007 Krakow, Poland
[4] Polish Acad Sci, Inst Nat Conservat, PL-00901 Warsaw, Poland
[5] Jagiellonian Univ, Inst Geog & Spatial Planning, PL-31007 Krakow, Poland