[1] Univ Oxford, Weatherall Inst Mol Med, Oxford OX1 2JD, England
[2] Univ Oxford, Dept Clin Neurol, Oxford OX1 2JD, England
[3] Cyprus Inst Neurol & Genet, Dept Clin Neurosci, Nicosia, Cyprus
[4] Univ Penn, Sch Med, Dept Neurol, Philadelphia, PA 19104 USA