[1] Natl Agr Res Ctr, Tsukuba, Ibaraki 3058666, Japan
[2] Osaka Univ, Inst Prot Res, Suita, Osaka 5650871, Japan
[3] Peking Univ, Coll Life Sci, Peking Yale Joint Ctr Plant Mol Genet & Agrobiote, Natl Lab Prot Engn & Plant Genet Engn, Beijing 100871, Peoples R China
[4] Univ Hyogo, Dept Life Sci, Akoh, Hyogo 6781297, Japan