[1] US Geol Survey, Pacific Coastal & Marine Sci Ctr, Santa Cruz, CA 95060 USA
[2] US Geol Survey, Forest & Rangeland Ecosyst Sci Ctr, Olymp Field Stn, Port Angeles, WA USA
[3] Natl Pk Serv, Olymp Natl Pk, Port Angeles, WA USA
[4] Bur Reclamat Sedimentat & River Hydraul Grp, Denver, CO USA
[5] NOAA, Natl Marine Fisheries Serv, Seattle, WA 98115 USA
[6] US Geol Survey, Washington Water Sci Ctr, Tacoma, WA USA
[7] US Geol Survey, Southwest Biol Sci Ctr, Flagstaff, AZ 86001 USA