[1] GEOMAR Helmholtz Ctr Ocean Res Kiel, D-24148 Kiel, Germany
[2] Univ Bremen, MARUM Ctr Marine Environm Sci, D-28359 Bremen, Germany
[3] Univ Bremen, Fac Geosci, D-28359 Bremen, Germany
[4] Univ Basel, Dept Environm Sci, CH-4056 Basel, Switzerland
[5] Univ Tromso, Dept Geol, CAGE Ctr Arctic Gas Hydrate Environm & Climate, N-9037 Tromso, Norway
[6] Natl Oceanog Ctr, Southampton SO14 3ZH, Hants, England