[1] IPP RAS, KA Timiryazev Inst Plant Physiol RAS, 35 Botanicheskaya St, Moscow 127276, Russia
[2] Russian Acad Sci, Papanin Inst Biol Inland Waters, Yaroslavl 152742, Russia
[3] Univ Colorado, Museum Nat Hist, Boulder, CO 80309 USA
[4] Univ Colorado, Dept Ecol & Evolutionary Biol, Boulder, CO 80309 USA