[1] Chinese Acad Sci, Xishuangbanna Trop Bot Garden, Key Lab Trop Plant Resource & Sustainable Use, Key Lab Trop Forest Ecol, Mengla 666303, Yunnan, Peoples R China
[2] Chinese Acad Sci, Kunming Inst Bot, Key Lab Biodivers & Biogeog, Kunming 650201, Yunnan, Peoples R China
[3] Royal Bot Garden Edinburgh, Edinburgh EH3 5LR, Midlothian, Scotland