[1] Univ So Queensland, Australian Ctr Sustainable Catchments, Toowoomba, Qld 4350, Australia
[2] Univ So Queensland, Fac Sci, Toowoomba, Qld 4350, Australia
[3] Australian Inst Hlth & Welf, Canberra, ACT 2601, Australia
[4] Goulburn Murray Water, Kerang, Vic 3579, Australia