[1] Natl Cheng Kung Univ, Dept Biotechnol & Bioind Sci, Tainan 701, Taiwan
[2] Natl Cheng Kung Univ, Int Ctr Sci Dev Shrimp Aquaculture, Tainan 701, Taiwan
[3] Tokyo Univ Marine Sci & Technol, Grad Sch Marine Sci & Technol, Tokyo 1088477, Japan
[4] Univ Santo Tomas, Dept Biol Sci, Manila 1015, Philippines
[5] Chulalongkorn Univ, Fac Sci, Dept Biochem, Bangkok 10330, Thailand
[6] Fisheries & Aquaculture Sci Cefas, Ctr Environm, Pathol & Mol Systemat Team, Weymouth DT4 8UB, England