[1] Shandong Univ Sci & Technol, Qingdao 266510, Peoples R China
[2] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Jiangsu, Peoples R China
[3] Russian Acad Sci, Inst Paleontol, Profsoyuznaya Ul 123, Moscow 117997, Russia
[4] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China