[1] De La Salle Univ, Br Alfred Shields FSC Ocean Res Ctr, Manila 1004, Philippines
[2] De La Salle Univ, Integrated Sch Res & Dev Off, Binan 4024, Laguna, Philippines
[3] Univ Philippines Diliman, Marine Environm & Resources Fdn, Inst Marine Sci, Quezon City 1101, Philippines
[4] Univ Philippines Diliman, Inst Marine Sci, Quezon City 1101, Philippines
[5] Ninoy Aquino Parks & Wildlife Ctr, Coastal & Marine Div, Biodivers Management Bur, Dept Environm & Nat Resources, Quezon City 1100, Philippines
[6] Univ Hawaii Manoa, Dept Geog, Honolulu, HI 96822 USA