[1] INRA, UMR 1229, F-21000 Dijon, France
[2] Univ Burgundy, UMR 1229, F-21000 Dijon, France
[3] Acad Sci Czech Republ, Inst Soil Biol, Ctr Biol, CR-37005 Ceske Budejovice, Czech Republic
[4] Univ S Bohemia, Fac Sci, Ceske Budejovice 37005, Czech Republic
[5] Ctr Rech Orleans, INRA, INFOSOL Unit, US 1106, F-45166 Olivet, France