[1] Chulalongkorn Univ, Fac Sci, Program Biotechnol, Bangkok 10330, Thailand
[2] Field Museum, Sci & Educ, 1400 S Lake Shore Dr, Chicago, IL 60605 USA
[3] ABL Herbarium, Gerrit van der Veenstr 107, NL-3762 XK Soest, Netherlands
[4] Chulalongkorn Univ, Fac Sci, Dept Bot, Bangkok 10330, Thailand
[5] Ramkhamhang Univ, Fac Sci, Dept Biol, Bangkok 10240, Thailand