[1] Grad Univ Adv Studies, Dept Evolutionary Studies Biosyst, Kanagawa 2400193, Japan
[2] Univ Tokyo, Dept Cognit & Behav Sci, Tokyo, Japan
[3] Aska Pharma Med, Kawasaki, Kanagawa, Japan
[4] Chimpanzee Sanctuary Uto, Kumamoto, Japan
[5] Univ Tokyo, Dept Anthropol, Lab Human Genet, Tokyo, Japan
[6] Univ Tokyo, Dept Biomed Sci, Tokyo, Japan