[1] Univ Aix Marseille 2, URMITE, UMR 6236, CNRS,IRD,Fac Med & Pharm, F-13385 Marseille 05, France
[2] Govt Lao PDR, NAFRI, Minist Agr & Forestry, Viangchan, Laos
[3] CSIRO Sustainable Ecosyst, Australian Natl Wildlife Collect, Canberra, ACT, Australia
[4] Mahosot Hosp, Wellcome Trust Mahosot Hosp Oxford Trop Med Res C, Microbiol Lab, Viangchan, Laos
[5] Univ Oxford, Ctr Trop Med, Churchill Hosp, Oxford OX1 2JD, England