[1] Kyoto Univ, Grad Sch Med, Kyoto 6068501, Japan
[2] Kyoto Univ, Inst Integrated Cell Mat Sci, Ctr iPS Cell Res & Applicat, Kyoto 6068501, Japan
[3] Kyoto Univ, Inst Chem Res, Kyoto 6068501, Japan
[4] Kyoto Univ, Inst Integrated Cell Mat Sci, Kyoto 6068501, Japan