[1] Boston Univ, Dept Geog & Environm, Boston, MA 02215 USA
[2] CSIRO Marine & Atmospher Resources, Canberra, ACT 2601, Australia
[3] CSIRO Sustainable Ecosyst, Clayton, Vic 3169, Australia
[4] Harvard Univ, Dept Organism & Evolutionary Biol, Cambridge, MA 02138 USA
[5] CUNY Hunter Coll, Dept Geog, New York, NY 10021 USA