[1] Ocean Univ China, Inst Evolut Marine Biodivers, 5 Yushan Rd, Qingdao 266003, Peoples R China
[2] Univ Florida, Dept Entomol & Nematol, Honey Bee Res & Extens Lab HBREL, Steinmetz Hall,Nat Area Dr,POB 110620, Gainesville, FL 32611 USA
[3] Heidelberg Univ, Inst Pharm & Mol Biotechnol IPMB, Neuenheimer Feld 364, D-69120 Heidelberg, Germany