[1] Univ Calif San Francisco, Gladstone Inst Neurol Dis, San Francisco, CA 94158 USA
[2] Univ Calif San Francisco, Chem & Chem Biol Grad Program, San Francisco, CA 94158 USA
[3] Univ Calif San Francisco, Med Scientist Training Program, San Francisco, CA 94158 USA
[4] Univ Calif San Francisco, Dept Biochem & Biophys, San Francisco, CA 94158 USA
[5] Univ Calif San Francisco, Dept Neurol, San Francisco, CA 94158 USA