[1] St Thomas Hosp, Dept Ophthalmol, London, England
[2] Kings Coll London, Dept Twin Res & Genet Epidemiol, London, England
[3] Maidstone Hlth Author, Dept Ophthalmol, Maidstone, Kent, England
[4] Kings Coll London, Dept Ophthalmol, St Thomas Hosp Campus,Westminster Bridge Rd, London SE1 7EH, England
[5] Univ Cambridge, Dept Physiol Dev & Neurosci, Cambridge, England
[6] Moorfield Eye Hosp NHS Fdn Trust, London, England
[7] UCL Inst Ophthalmol, London, England