[1] Sun Yat Sen Univ, Museum Biol, State Key Lab Biocontrol, Sch Ecol,Sch Life Sci, Guangzhou 510275, Guangdong, Peoples R China
[2] Nat Hist Museum Guangxi, Nanning 530012, Guangxi, Peoples R China
[3] Guangdong Shimentai Natl Nat Reserve, Qingyuan 513000, Guangdong, Peoples R China
[4] La Sierra Univ, Dept Biol, Herpetol Lab, Riverside, CA 92515 USA
[5] San Diego Nat Hist Museum, Dept Herpetol, POB 121390, San Diego, CA 92112 USA