[1] Seoul Natl Univ, Dept Vet Med, Virol Lab, Coll Vet Med, Seoul 151742, South Korea
[2] Seoul Natl Univ, Program Vet Sci BK21, Seoul 151742, South Korea
[3] Natl Vet Res & Quarantine Serv, Anyang 4308, Gyeonggi Do, South Korea
[4] Anim Genet Inc, Suwon 443823, South Korea
[5] Green Cross Vet Prod Co Ltd, Res Unit, Yongin 449903, South Korea