Yield, yield attributes and photosynthetic physiological characteristics of dryland wheat (Triticum aestivum L.)/maize (Zea mays L.) strip intercropping

Intercropping has been widely adopted by farmers for higher yield compared to monoculture, and the border effect was responsible for the yield advantage in irrigated areas. However, few studies have investigated the border effect based on photosynthesis, especially under rainfed conditions. Here, we evaluated the yield in rainfed wheat/maize strip relay intercropping, and explored the associated differences in yield components and physiological process compared to sole crops. A two-year field experiments was conducted including three treatments (sole wheat, sole maize and wheat/maize intercropping) in Yangling, located in the semi-humid region of northwest China. Grain yield, yield components, chlorophyll relative content (SPAD) and photosynthetic parameters in different rows were measured for wheat and maize. Results showed that wheat/maize intercropping increased the land use efficiency and total yield of wheat and maize under rainfed conditions. The yield of intercropped wheat was significantly (p < 0.05) increased by 35% on average over two years, resulting not only from the first border rows but also from the second border rows. The yield improvement in the first border rows relative to sole wheat was attributed to the increase of ear number (56%, p < 0.05), kernel number per ear (14%, p < 0.05) and thousand kernel weight (12%, p < 0.05), but the yield improvement in the second border rows was only attributed to the ear number (22%, p < 0.05). The yield of intercropped maize was not significantly (p > 0.05) decreased (6%), mainly attributed to the border rows (19% lower than in sole maize), in which the kernel number per cob and thousand kernel weight were 14% and 8% (p < 0.05) lower than that in sole maize. This was because that the SPAD and photosynthetic rate of maize in intercropping was suppressed during the co-growth period (about 62 days). Although the two had a partly growth recovery after wheat harvest, the growth recovery was not complete, and this was responsible for the reduction of maize yield in intercropping. This study demonstrates that the importance of yield components and the photosynthesis basis for yield advantage in intercropping systems, and reducing the adverse effect of dominate crop on subordinate crops could better exert the advantages of intercropping.