[1] Chinese Acad Sci, South China Sea Inst Oceanol, State Key Lab Trop Oceanog, 164 West Xingang Rd, Guangzhou 510301, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab Guangzho, Guangzhou 511458, Peoples R China
[3] Chinese Acad Sci, Inst South China Sea Ecol & Environm Engn, Guangzhou 510301, Peoples R China
[4] Univ Chinese Acad Sci, Beijing 100049, Peoples R China