[1] Chinese Acad Sci, Chengdu Inst Biol, Chengdu 610041, Peoples R China
[2] Northwest Agr & Forest Univ, Coll Forestry, Lab Ecol Forecasting & Global Change, Yangling 712100, Peoples R China
[3] Chinese Acad Sci, Zoige Peatland & Global Change Res Stn, Hong yuan 624400, Peoples R China
[4] Int Ctr Integrated Mt Dev, Kathmandu, Nepal
[5] Univ Quebec, Inst Environm Sci, Dept Biol Sci, Montreal, PQ C3H 3P8, Canada
[6] Chinese Acad Sci, Inst Mt Hazards & Environm, Chengdu 610041, Peoples R China
[7] Mem Univ Newfoundland, Sustainable Resource Management, Corner Brook, NL A2H 6P9, Canada
[8] Chinese Acad Sci, Inst Microbiol, Beijing 100101, Peoples R China
[9] Chinese Acad Sci, Northwest Inst Plateau Biol, Xining 810008, Peoples R China
[10] Southwest Univ Sci & Technol, Life Sci & Engn Coll, Mianyang 621010, Peoples R China
[11] Univ Oklahoma, Ctr Spatial Anal, Dept Microbiol & Plant Biol, Norman, OK 73019 USA