[1] Univ Ouagadougou, Lab Biol & Ecol Vegetales, Unite Format & Rech Sci Vie & Terre, Ouagadougou 03, Burkina Faso
[2] Univ Ouagadougou, Lab Biochim & Chim Appl, Unite Format & Rech Sci Vie & Terre, Ouagadougou 03, Burkina Faso
[3] Univ Gand, Fac Sci Bioingn, Lab Agron Trop & Subtrop & Ethnobot, B-9000 Ghent, Belgium