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Univ Fed Rio Grande Do Sul, Dept Zool, Lab Entomol Sistemat, Inst Biociencias, Ave Bento Goncalves 9500, Porto Alegre, Rio Grande do S, Brazil Univ Fed Rio Grande Do Sul, Dept Zool, Lab Entomol Sistemat, Inst Biociencias, Ave Bento Goncalves 9500, Porto Alegre, Rio Grande do S, Brazil