[1] Osaka Univ, Grad Sch Med, Dept Neurosci & Cell Biol, Osaka, Japan
[2] Osaka Psychiat Med Ctr, Osaka Psychiat Res Ctr, Addict Res Unit, Osaka, Japan
[3] Osaka Univ, Grad Sch Med, Dept Neuromed Sci, Osaka, Japan
[4] Osaka Univ, Grad Sch Med, Dept Mol Neurosci, Osaka, Japan
[5] Kanazawa Univ, Grad Sch Med Sci, Dept Neuroanat, Kanazawa, Ishikawa, Japan
[6] Osaka Univ, United Grad Sch Child Dev, Dept Child Dev & Mol Brain Sci, Osaka, Japan