[1] US Geol Survey, Great Lakes Sci Ctr, Hammond Bay Biol Stn, 11188 Ray Rd, Millersburg, MI 49759 USA
[2] Michigan State Univ, Ctr Syst Integrat & Sustainabil, 1405 South Harrison Rd,115 Manly Miles Bldg, E Lansing, MI 48823 USA
[3] Great Lakes Fishery Commiss, 2100 Commonwealth Blvd,Suite 100, Ann Arbor, MI 48105 USA
[4] Washington Dept Fish & Wildlife, Div Sci, Wild Salmonid Prod Evaluat Unit, Fish Program, 1111 Washington St Southeast, Olympia, WA 98501 USA