[1] Res Org Informat & Syst, Div Human Genet, Dept Integrated Genet, Natl Inst Genet, Mishima, Shizuoka 4118540, Japan
[2] Grad Univ Adv Studies, Dept Genet, Sch Life Sci, Mishima, Shizuoka 4118540, Japan
[3] Japan Sci & Technol Agcy, PRESTO, Kawaguchi 3320012, Japan
[4] RIKEN, Ctr Dev Biol, Lab Mammalian Epigenet Studies, Kobe, Hyogo 6500047, Japan
[5] Harvard Univ, Sch Med, Massachusetts Gen Hosp, Cardiovasc Res Ctr, Charlestown, MA 02139 USA