[1] Russian Acad Sci, All Russian Inst Plant Protect, Shosse Podbelskogo 3, St Petersburg 196608, Russia
[2] Natl Acad Sci Ukraine, Schmalhausen Inst Zool, B Khmelnitskogo 15, UA-01601 Kiev, Ukraine
[3] Russian Acad Sci, AA Borissiak Paleontol Inst, Profsoyuznaya Str 123, Moscow 117647, Russia
[4] Natl Museum Scotland, Dept Nat Sci, Chambers St, Edinburgh EH1 1JF, Midlothian, Scotland
[5] Univ Bonn, Inst Geosci, D-53115 Bonn, Germany
[6] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Peoples R China