[1] Univ Cape Town, Nansen Tutu Ctr Marine Environm Res, Cape Town, South Africa
[2] Univ Cape Town, Marine Res Inst, Dept Oceanog, Cape Town, South Africa
[3] Inst Marine Res, Bergen, Norway
[4] Nelson Mandela Univ, Reg Ocean Sci & Marine Food Secur Res Ctr, Ocean Sci Campus, Port Elizabeth, South Africa