[1] Chinese Acad Sci, Innovat Acad Seed Design, Wuhan Bot Garden, CAS Key Lab Plant Germplasm Enhancement & Special, Wuhan 430074, Peoples R China
[2] Hubei Hongshan Lab, Wuhan 430070, Peoples R China
[3] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[4] Zhejiang Univ, Zhejiang Prov Key Lab Hort Plant Integrat Biol, Hangzhou 310058, Peoples R China
[5] Cornell Univ, Boyce Thompson Inst, Ithaca, NY 14853 USA
[6] USDA ARS, Robert W Holley Ctr Agr & Hlth, Ithaca, NY 14853 USA