[1] Ocean Univ China, Phys Oceanog Lab, Qingdao, Peoples R China
[2] Ocean Univ China, Coll Ocean & Atmospher Sci, Qingdao, Peoples R China
[3] Ocean Univ China, Key Lab Marine Environm & Ecol, Minist Educ, Qingdao, Peoples R China
[4] Bedford Inst Oceanog Fisheries & Oceans Canada, Ocean & Ecosyst Sci Div, Dartmouth, NS, Canada