[1] Univ Bordeaux, CNRS, PACEA UMR5199, Allee Geoffroy St Hilaire,CS 50023, F-33615 Pessac, France
[2] Inst Francais Afrique Sud, 62 Juta St, Johannesburg, South Africa
[3] Univ Witwatersrand, Sch Geog Archaeol & Environm Studies, Johannesburg, South Africa
[4] Univ Witwatersrand, Evolutionary Studies Inst, Johannesburg, South Africa
[5] Univ Bordeaux Montaigne, CNRS, IRAMAT UMR5060, Pessac, France