[1] Univ Montana, Wildlife Biol Program, Dept Ecosyst & Conservat Sci, WA Franke Coll Forestry & Conservat, 32 Campus Dr, Missoula, MT 59812 USA
[2] Alberta Conservat Assoc, 817 4th Ave South 400, Lethbridge, AB T1J 0P3, Canada
[3] Natl Wildlife Federat, Prairies & Pacific Pacif Reg, Northern Rockies, 240 North Higgins Ave,Suite 2, Missoula, MT 59802 USA