[1] RIKEN, Tsukuba Inst, Plant Mol Biol Lab, Tsukuba, Ibaraki 3050074, Japan
[2] RIKEN, Genom Sci Ctr, Plant Mutation Explorat Team, Plant Funct Genom Res Grp, Yokohama, Kanagawa 2300045, Japan
[3] Minist Agr Forestry & Fisheries, Japan Int Res Ctr Agr Sci, Biol Resources Div, Tsukuba, Ibaraki 3050851, Japan