[1] Univ Quebec, Inst Sci Mer Rimouski, Rimouski, PQ G5L 3A1, Canada
[2] Univ Sherbrooke, Dept Phys, Sherbrooke, PQ J1K 2R1, Canada
[3] Univ Laval, Dept Biol, Quebec City, PQ G1V 0A6, Canada
[4] Fisheries & Oceans Canada, Maurice Lamontagne Inst, Mont Joli, PQ G5H 3Z4, Canada
[5] Ctr Eau Terre Environm, Quebec City, PQ G1K 9A9, Canada