[1] Univ Calgary, Hotchkiss Brain Inst, Dept Cell Biol & Anat, Calgary, AB T2N 4N1, Canada
[2] Univ Calgary, Hotchkiss Brain Inst, Dept Psychiat, Calgary, AB T2N 4N1, Canada
[3] Univ Massachusetts Boston, Dept Psychol, Boston, MA 02125 USA
[4] Univ South Carolina, Dept Pharmacol Physiol & Neurosci, Columbia, SC 29208 USA
[5] WJB Dorn VA Med Ctr, Columbia, SC 29208 USA