[1] Ctr Environm Fisheries & Aquaculture Sci, Weymouth Lab, Weymouth DT4 8UB, Dorset, England
[2] Univ Oslo, Dept Biol, N-0316 Oslo, Norway
[3] Norwegian Inst Water Res NIVA, N-0349 Oslo, Norway
[4] Marine Scotland, Marine Lab, Aberdeen AB11 9DB, Scotland
[5] Deltares, Marine & Coastal Syst, NL-2600 MH Delft, Netherlands
[6] Vrije Univ Amsterdam, Inst Environm Studies, NL-1081 HV Amsterdam, Netherlands