[1] Taizhou Univ, Sch Life Sci, Zhejiang Prov Key Lab Plant Evolutionary & Conser, 1139 Shifu Rd, Jiaojiang Dist 318000, Taizhou, Peoples R China
[2] Huaibei Normal Univ, Sch Life Sci, Anhui Key Lab Resource & Plant Biol, 100 Dongshan Ave, Xiangshan Dist 235000, Huaibei, Peoples R China
[3] Chinese Acad Agr Sci, Inst Environm & Sustainble Dev Agr, 12 Zhongguancun South St, Beijing 100081, Peoples R China
[4] Univ Western Australia, UWA Sch Agr & Environm, 35 Stirling Highway, Crawley, WA 6009, Australia
[5] Chinese Acad Agr Sci, Inst Agr Resources & Reg Planning, 12 Zhongguancun South St, Beijing 100081, Peoples R China