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Sun Yat Sen Univ, Coll Ecol & Evolut, Guangdong Higher Educ Inst, Key Lab Biodivers Dynam & Conservat,State Key Lab, Guangzhou 510275, Guangdong, Peoples R China
CSIRO, Natl Res Collect Australia, Australian Natl Insect Collect, Canberra, ACT 2601, Australia Sun Yat Sen Univ, Coll Ecol & Evolut, Guangdong Higher Educ Inst, Key Lab Biodivers Dynam & Conservat,State Key Lab, Guangzhou 510275, Guangdong, Peoples R China