[1] Austin Hlth, Gastroenterol, Heidelberg, Vic, Australia
[2] St Vincents Hosp, Immunol Res Ctr, Fitzroy, Vic, Australia
[3] Univ Melbourne, Austin Acad Ctr, Med, Heidelberg, Vic, Australia
[4] Univ Melbourne, Dept Med Dent & Hlth Sci, Parkville, Vic, Australia
[5] Univ Melbourne, Peter Doherty Inst Infect & Immun, Dept Microbiol & Immunol, Parkville, Vic, Australia