[1] Yale Univ, Yale Sch Environm, New Haven, CT 06520 USA
[2] New York Bot Garden, Bronx, NY 10458 USA
[3] Smithsonian Trop Res Inst, Balboa, Panama
[4] Univ Rosario, Fac Nat Sci, Dept Biol, Bogota, Colombia
[5] Marquette Univ, Dept Biol Sci, Milwaukee, WI USA