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Chinese Acad Agr Sci, Inst Vegetables & Flowers, Key Lab Biol & Genet Improvement Hort Crops, Minist Agr, 12 Zhong Guan Cun Nandajie St, Beijing 100081, Peoples R China Chinese Acad Agr Sci, Inst Vegetables & Flowers, Key Lab Biol & Genet Improvement Hort Crops, Minist Agr, 12 Zhong Guan Cun Nandajie St, Beijing 100081, Peoples R China